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In Chicago, Wesley Gibson works at a dead-end desk job with an overbearing boss, takes medication for panic attacks, and lives with his abrasive girlfriend Cathy who cheats on him with his co-worker and best friend, Barry. One evening, Wesley is told by a woman named Fox that his recently murdered father was an assassin, and the killer, Cross, is now hunting him. When Cross and Fox engage in a shootout, Wesley panics and is pursued by Cross. Fox flips Wesley into her car and then executes an escape. Wesley awakens in a factory surrounded by Fox and other assassins. The group's leader, Mr. Sloan, forces Wesley at gunpoint to shoot the wings off of several flies, which he does, much to Wesley's shock. Sloan explains that Wesley's panic attacks are actually a rare ability that allows him to produce massive amounts of adrenaline, slowing his perception of his surroundings and granting him superhuman strength and speed. He reveals that Wesley's father, as well as Cross, were members of the Fraternity, a society of assassins that maintains balance in the world. Sloan wants to train him so that he may help kill Cross.

A panicked Wesley leaves the building and awakens the next morning in his apartment. He discovers that his bank account now contains $3.6 million. Filled with new confidence, he insults his boss in front of the whole office and hits Barry in the face with a keyboard. Fox, waiting for Wesley, chauffeurs him to the Fraternity headquarters, a repurposed textile mill. Wesley trains under the Fraternity's cruel tutelage, learning to control his abilities; Fox teaching him speed, the Gunsmith teaching him marksmanship, such as curving a bullet, the Butcher teaching him hand-to-hand combat, the Exterminator teaching him explosives, and the Repairman teaching him pain resistance.